Boraginaceae s.str.
This list of genera was made in April 2024 according to Plants of the World Online, Stevens 2001 onwards (APWeb), Chacón et al., 2016, Khoshsokhan-Mozaffar et al., 2018, Luebert et al., 2016, Nasim et al., 2017 and Selvi et al., 2017. This list includes only genera present in our herbaria: M and MSB.
| All genera (accepted genera in bold) | Stored under | Synonyms included |
|---|---|---|
| Actinocarya Benth. | Microula Benth. | |
| Adelinia J.I.Cohen | ||
| Aegonychon Gray (prev. under Lithospermum L.) | ||
| Afrotysonia Rauschert | ||
| Alkanna Tausch | ||
| Allocarya Greene | Plagiobothrys Fisch. & C.A.Mey. | |
| Amsinckia Lehm. | ||
| Anchusa L. | ||
| Anoplocaryum Ledeb. | ||
| Antiotrema Hand.-Mazz. | ||
| Antiphytum DC. ex Meisn. | ||
| Arnebia Forssk. | Macrotomia DC. | |
| Asperugo L. | ||
| Borago L. | ||
| Bothriospermum Bunge | ||
| Brachybotrys Maxim. ex Oliv. | ||
| Brunnera Steven | ||
| Buglossoides Moench | ||
| Caccinia Savi | ||
| Cerinthe L. | ||
| Craniospermum Lehm. | ||
| Cryptantha Lehm. ex G.Don | Eremocarya Greene, Greeneocharis Gürke & Harms | |
| Cynoglossum L. | Ivanjohnstonia Kazmi, Kuschakewiczia Regel & M.Smirn., Lindelofia Lehm., Mattiastrum (Boiss.) Brand, Paracaryum Boiss, Pardoglossum Barbier & Mathez, Rindera Pall., Solenanthus Ledeb., Trachelanthus Kunze | |
| Cynoglottis (Gușul.) Vural & Kit Tan | ||
| Cystostemon Balf.f. | Vaupelia Brand | |
| Decalepidanthus Riedl (prev. under Pseudomertensia Riedl) | Pseudomertensia Riedl | |
| Echiochilon Desf. | Sericostoma Stocks | |
| Echioides Ortega | Huynhia Greuter | |
| Echiostachys Levyns | ||
| Echium L. | ||
| Elizaldia Willk. | Nonea Medik. | |
| Eremocarya Greene | Cryptantha Lehm. ex G.Don | |
| Eritrichium Schrad. ex Gaudin | ||
| Gastrocotyle Bunge | ||
| Glandora D.C.Thomas, Weigend & Hilger | ||
| Greeneocharis Gürke & Harms | Cryptantha Lehm. ex G.Don | |
| Hackelia Opiz (prev under Lappula Moench.) | ||
| Halacsya Dörfl. | ||
| Harpagonella A.Gray | ||
| Havilandia Stapf | Trigonotis Steven | |
| Heterocaryum A.DC. | ||
| Huynhia Greuter (prev. under Echinoides Ortega) | Echioides Ortega | |
| Iberodes M.Serrano, R.Carbajal & S.Ortiz | ||
| Ivanjohnstonia Kazmi | Cynoglossum L. | |
| Kuschakewiczia Regel & M.Smirn. | Cynoglossum L. | |
| Lappula Moench | Omphalolappula Brand, Lepechiniella Popov, Sclerocaryopsis Brand | |
| Lasiocaryum I.M.Johnst. | ||
| Lepechiniella Popov | Lappula Moench | |
| Lindelofia Lehm. | Cynoglossum L. | |
| Lithodora Griseb. | ||
| Lithospermum L. | Macromeria D.Don, Onosmodium Michx., Psilolaemus I.M.Johnst. | |
| Lobostemon Lehm. | ||
| Lycopsis L. (prev. under Anchusa L.) | ||
| Macromeria D.Don | Lithospermum L. | |
| Macrotomia DC. | Arnebia Forssk. | |
| Maharanga DC. (prev. under Onosma L.) | ||
| Mairetis I.M.Johnst. | ||
| Mattiastrum (Boiss.) Brand | Cynoglossum L. | |
| Megacaryon Boiss. | ||
| Megastoma Bonnet & Baratte | Ogastemma Brummitt | |
| Mertensia Roth | ||
| Microcaryum I.M.Johnst. | ||
| Microparacaryum (Popov ex Riedl) Hilger & Podlech | ||
| Microula Benth. | Actinocarya Benth. | |
| Mimophytum Greenm. | ||
| Moltkia Lehm. | ||
| Moltkiopsis I.M.Johnst. | ||
| Moritzia DC. ex Meisn. | ||
| Myosotidium Hook. | ||
| Myosotis L. | Trigonocaryum Trautv. | |
| Neatostema I.M.Johnst. | ||
| Nesocaryum I.M.Johnst. | ||
| Nihon A.Otero, Jim.Mejías, Valcárcel & P.Vargas | ||
| Nonea Medik. | Elizaldia Willk. | |
| Ogastemma Brummitt | Megastoma Bonnet & Baratte | |
| Omphalodes Mill. | ||
| Omphalolappula Brand | Lappula Moench | |
| Onosma L. | ||
| Onosmodium Michx. | Lithospermum L. | |
| Oreocarya Greene | Cryptantha Lehm. ex G.Don | |
| Paracaryum Boiss. | Cynoglossum L. | |
| Paraskevia W.Sauer & G.Sauer | Pulmonaria L. | |
| Pardoglossum Barbier & Mathez | Cynoglossum L. | |
| Pectocarya DC. ex Meisn. | ||
| Pentaglottis Tausch | ||
| Phyllocara Guşul. | ||
| Plagiobothrys Fisch. & C.A.Mey. | Allocarya Greene | |
| Pontechium Böhle & Hilger | ||
| Procopiania Gușul. | Symphytum L. | |
| Pseudoheterocaryum Kaz.Osaloo & Saadati | ||
| Pseudolappula Khoshsokhan & Kaz.Osaloo (prev. under Lappula sinaica (A.DC.) Asch. & Schweinf.) | ||
| Pseudomertensia Riedl | Decalepidanthus Riedl | |
| Psilolaemus I.M.Johnst. | Lithospermum L. | |
| Pulmonaria L. | Paraskevia W.Sauer & G.Sauer | |
| Rindera Pall. | Cynoglossum L. | |
| Rochelia Rchb. | ||
| Sclerocaryopsis Brand | Lappula Moench | |
| Selkirkia Hemsl. | ||
| Sericostoma Stocks | Echiochilon Desf. | |
| Solenanthus Ledeb. | Cynoglossum L. | |
| Stenosolenium Turcz. (prev. under Arnebia Forsk.) | ||
| Suchtelenia Kar. ex Meisn. | ||
| Symphytum L. | Procopiania Guşul. | |
| Thaumatocaryon Baill. (prev. under Antiphytum DC) | ||
| Thyrocarpus Hance | ||
| Trachelanthus Kunze | Cynoglossum L. | |
| Trachystemon D.Don | ||
| Trichodesma R.Br. | ||
| Trigonocaryum Trautv. | Myosotis L. | |
| Trigonotis Steven | Havilandia Stapf | |
| Vaupelia Brand | Cystostemon Balf.f. |
References
- Chacón, J., Luebert, F., Hilger, H.H., Ovchinnikova, S., Selvi, F., Cecchi, L., Guilliams, C.M., Hasenstab-Lehman, K., Sutorý, K., Simpson, M.G. and Weigend, M. (2016). The borage family (Boraginaceae s.str.): A revised infrafamilial classification based on new phylogenetic evidence, with emphasis on the placement of some enigmatic genera. Taxon, 65: 523-546. https://doi.org/10.12705/653.6
- Luebert, F., Cecchi, L., Frohlich, M.W., Gottschling, M., Guilliams, C.M., Hasenstab-Lehman, K.E., Hilger, H.H., Miller, J.S., Mittelbach, M., Nazaire, M., Nepi, M., Nocentini, D., Ober, D., Olmstead, R.G., Selvi, F., Simpson, M.G., Sutorý, K., Valdés, B., Walden, G.K. and Weigend, M. (2016). Familial classification of the Boraginales. Taxon, 65: 502-522. https://doi.org/10.12705/653.5
- Khoshsokhan-Mozaffar, M., Sherafati, M., Kazempour-Osaloo, S. (2018). Molecular phylogeny of the tribe Rochelieae (Boraginaceae, Cynoglossoideae) with special reference to Lappula. Annales Botanici Fennici, 55(4-6), 293-308. https://doi.org/10.5735/085.055.0411
- Nasim, S., Khoshsokhan-Mozaffar, M., Mahboubeh, S., Kazempour-Osaloo, S. (2017). Pseudoheterocaryum, a new genus segregated from Heterocaryum (Boraginaceae) on the basis of molecular data. Australian Systematic Botany 30, 105-111. https://doi.org/10.1071/SB16022
- Selvi, F., Cecchi, L., Hilger, H. H., & Coppi, A. (2017). A reappraisal of the genus Megacaryon (Boraginaceae, Lithospermeae) based on molecular, morphological, and karyological evidence. Systematics and Biodiversity, 15(6), 552–563. https://doi.org/10.1080/14772000.2017.1290707
- Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 14, July 2017 [and more or less continuously updated since].
Foto: Myosotis L. © Luo Chen